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CALCULATING SURFACE AREA 829 Calculating …

Landscape area is almost always presented interms of planimetric area , as if a square kilometer ina mountainous area represents the same amount ofland area as a square kilometer in the of home ranges for wildlife speciesgenerally use planimetric area even when describ-ing mountain goats (Oreamnos americanus) andpumas (Felis concolor). But if a species behaviorand population dynamics are functions of availableresources, and if those resources are spatially limit-ed, I suggest assessing resources using SURFACE areaof the area also is a basis for a useful measure oflandscape topographic roughness. The SURFACE -arearatio of any particular region on the landscape canbe calculated by dividing the SURFACE area of thatregion by the planimetric example,Bowden et al.

points P and Q: where a=planimetric (horizontal) distance from P to Q, b=difference in elevation between P and Q, c=surface distance from P to Q. Distance “b”is easy to calculate because it is sim-ply the absolute difference between the …

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